In the past, it was believed,there was many differences between humans and animals but now that list is shrinking.
Primates and non primate species can construct and use tools. McGrew 1992.Non human species have a sense of self. Gallup 1970.Presence of theory of mind. Individual knows that other individuals have different information than one self. Tomasello and Call 1997.Other species can communicate. Like Vervet monkeys. And can also fall into depression. Jane Goodall 1986.
Material culture. Chimpanzees constructing termite sticks. Goodall 1986. Chimpanzees also use hammer. use leaves to clean themselves with water.
West African ales used nut hammering whereas East African apes use leaves. This variation across chimpanzee populations regarding tool used and construction was found to constitut Culture by Whiten 1999.
Social custom. In Tanzania there is unique occurrence of "hand-clasp" grooming - where is animal raises and ARM grasping hands with the other. Different groups showing different technique. Nakamura 2004.
Raindance Sushil custom reported by Jane goodall 1986. not biological because it is only seen in East African populations and not in West Africa.